When the University of Alabama played Georgia in the 1964 football-season opener in Tuscaloosa, Archie Wade, Joffre Whisenton and Nathaniel Howard had seats next to the home team’s marching band.

Archie, a 55-year-old southern white rhino at the Jacksonville Zoo and Gardens, has died. He was the oldest male of his species in human care and a founder of the North American rhino population.